( 2017), while neither type-fixation nor diagnosis was given in Wang et al. However, this name is unfortunately not available taxonomically in the light of the International Code of Zoological Nomenclature (4th edition available at: ), because no Zoobank ID was given in Wang et al. Subsequently, the Southern Taiwan clade was split from the combined Central+Northern Ryukyu, and Northern Taiwan+Southern Ryukyu clades as a distinct species Buergeria otai by Wang et al. Genetic distances found among these three clades (12.8–17.3% in cytochrome b and 5.2–6.9% in 16S rRNA) are regarded as quite large compared with reported values among various congeneric anuran species, and each of these clades was thought to represent a taxonomically independent species ( Fig. ( 2015) studied the phylogeography of this species across its entire distributional range, and recognized three major clades in it: (1) the Southern Taiwan, (2) the Northern Taiwan+Southern Ryukyu, and (3) the Central+Northern Ryukyu clades. Buergeria japonica (Hallowell, 1861) was long considered a single, wide-ranging species occurring from the Tokara Islands of the Northern Ryukyus through the Central and the Southern Ryukyus, Japan, to Taiwan ( Okada, 1931 Nakamura and Uéno, 1963 Maeda and Matsui, 1999).
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